In one case outlined in medical histories obtained by the Globe, a 4-year-old monkey died of dehydration because she had no water spout in her cage. In another case, in 2010, the body of a monkey was found in a cage after it went through a heated power washer.
Harvard Medical School was fined more than $24,000 by the USDA for the center's violations of the Animal Welfare Act in 2013.
The Primate Research Center, which is home to more than 2,000 monkeys used in lab research, has been funded annually by more than $25 million in taxpayer money. While it is set to close at the end of May, the Globe's report raises serious questions about the normal conditions primates are kept in when used in research facilities.
Harvard responded to the Globe's inquiries with a statement outlining the changes it has made to the center since 2011.
To read the Globe's full report, click here.